A new paleobiogeographic subdivision of the Perigondwana terranes is based on the study of Upper Devonian and Lower Carboniferous calcareous foraminifers. The Brunovistulian, Moesian and Zonguldak terranes in Central and SE Europe and Asia Minor (the Brunovistulian group of terranes) show close paleobiogeographic relationships to the East European foraminiferal association. Consequently, they are regarded as a part of the Fennosarmatian Province which represented a centre for the diversification for most groups of Upper Devonian and Lower Carboniferous calcareous foraminifers. A lower degree of similarity with the East European Platform can be distinguished in the East Avalonian foraminiferal fauna while that of the Armorican group of terranes displays distinct differences constituting the separate Armorican Province. The results obtained support the idea that the Brunovistulian group of terranes formed the southern margin of Laurussia which was colliding with the Armorican group of terranes during the Variscan orogeny. In this respect, Brunovistulian group of terranes held a geotectonic position similar to that of East Avalonia even though the accretionary history of the terranes was different.